Education
•1982 B. S. in Fisheries, Tokyo University of Fisheries
•1984 M.S. in Fisheries, Tokyo University of Fisheries. Thesis: "Temporal and spatial dynamics of the zooplankton community in Ogochi Reservoir, Tokyo", major professor: M. Murano.
•1988 Ph. D. in Biology, Tokyo Metropolitan University. Thesis: "Demographical analysis on the life history traits of Daphnia galeata Sars in an artificial reservoir", major professor: Y. Watanabe.
Professional experience
•1987-1989 Officer, Education Bureau of Chiba Prefecture.
•1989-1992 Research Scientist, Department of Ecology, Natural History Museum and Institute, Chiba.
•1992-1994 Assistant Professor, Department of Biology, Tokyo Metropolitan University.
•1994-1995 Visiting Scientist, Department of Ecology, Evolution and Behavior, University of Minnesota.
•1995 - 2003 Associate Professor, Center for Ecological Research, Kyoto University.
•2003-present Professor, Div. Ecology and Evolutionary Biology, Graduate School of Life Sciences, Tohoku University