4.3. Floral organ sequence initiation
In Apiaceae, the diverse patterns of floral development have reported. The strict centripetal sequence of organ initiation has only found in Saniculoideae (Erbar and Leins 1985). By so far (~42 species), the diverse patterns of floral sequences have known in this family including centripetal, bidirectional, centrifugal with stamen dominance or sectorial group-like initiating (Payer, 1853, Erbar and Leins 1985, Jochmann 1854, Schuchardt 1881, Schumann 1890, Sieler 1870, Jurica 1922, Borthwick et al. 1931, Sattler 1973, Ajani et al 2016, Baczyński et al. 2022). The sectorial group-like development observed inSeseli ghafoorianum (Akhani) Pimenov and Kljuykov, Da. carota , and Orlaya grandiflora (L.) Hoffm. In these species, two common primordia fractionated alternatively in opposite sides, each splitting into sepal, petals and stamen (Ajani et al. 2016). Recently, this model observed in staminate flowers of Apiaceae with pseudanthia inflorescences, but studied only in peripheral staminate flowers (Baczyński et al. 2022). The similar group-like pattern also found inDi. persica . However, petals initiated as clockwise direction. In staminate flowers, furthermore, sepals developed late after elongation of petals and stamens. In perfect flowers, despite sepal suppression, petals and stamens resembled those organs in staminate flowers in sequencing of the organs. The interval time between initiation and elongation of the sepals filled out by fast synchronization of those organs in perfect flowers. In mature flowers, therefore, all flowers seems synchronized.