AUTHOR CONTRIBUTIONS
Shuai Li, Xinyang Tan, Zhen He and Rui Ji conceived the project, performed the experiments, analyzed the data, and wrote the manuscript. Rui Ji, Jichao Fang, Ary A. Hoffmann, and Chunqing Zhao supervised the project and edited the English language. Yali Li, Liu Yang and Lei Jiang helped in the sequencing data handling and arrangement. All authors read and approved the final manuscript.