Author Contributions

MA, AE, KMM, JP, HS, PW, CW, and BZ secured funding. GA, BZ, CW, HS, AE, and PW conceived the idea of the study. GA led and conducted fieldwork with the help of Per Larsson, KN, EM, EF and BZ. GA compiled and analysed the data and led the writing of the manuscript. MA created the script to estimate rMCP and CM contributed to the survival analysis. All authors contributed critically to the drafts and gave final approval for publication.