Table 2: Soil physical, chemical, and microbiological properties in each type of land cover. Mean values with standard errors in parentheses. Different letters indicate significant differences between land cover types (significant P-values in bold). BD: bulk density, WC: water content; WHC : water holding capacity; MWD : mean weight diameter; OC : organic carbon; TC: Total carbon, TN: total nitrogen; BR : basal respiration; MB : microbial biomass; qCO2 : metabolic quotient; FDAse : Fluorescein diacetate hydrolase.