AUTHOR CONTRIBUTIONS
A.M.Y., N.G.R., B.C.E., and D.P. conceived and designed the study. N.G.R, A.M.Y and D.P performed fieldwork, N.G.R. performed laboratory work, processed metabarcoding data and made all analyses. P.A., C.A., and B.C.E. supervised analyses and contributed to the discussion. Manuscript writing was led by N.G.R and A.M.Y., with contributions from all authors. All authors approved the final version of the manuscript.