Author contributions
Z.-M.H. and Z.Z. conceived the project. Z.-M.H. and J.Z. collected molecular data, J.Z. and P.F. performed genetic analyses, Z.Z. collected species distribution data and conducted ecological niche analyses with critical suggestions from J.M.K. and S.M. Z.-M.H., Z.Z., S.M., J.M.K., Z.-Q.Z., S.D., J.A., and A.J. interpreted and discussed the results, wrote and/or revised the manuscript. All authors approved the final version of the manuscript.