Acknowledgments
This study was funded by Agence Nationale de la Recherche (ANR-13-ECOT-0002 aquaDNA). GFF is partially funded by the European Research Council under the European CommunityHorizon 2020 Programme, Grant Agreement no. 772284 (IceCommunities). We thank SPYGEN staff for its help in laboratory, and S. Jolivet, M. Brulin, J. La Doaré, G. Coppa, M. Ferran and OPIE volunteers for the collection of reference specimens”.