Author contributions
J.Z., S.I., W.M. proposed the frost damage investigation. J.Z. visually scored the frost damage, implemented the molecular work and field work, analyzed the data and wrote the paper. B.B. supervised frost damage phenotyping; W.M. and JEM. contributed to the substantial paper writing and revisions. J.K. provided surveyed the parental lines and provided Bethlehem and CSL seeds for DH population construction; K.C. designed the field trials and statistically adjusted phenotype data; D.S., guided the group's perception about frost and helped interpreting the results;  Z.A., M.A., R.Y., participated in the DH population construction; J.Z., Y.Z., H.L., M.D., S.A., N.R., R.M., constructed the genetic linkage map; J.Z., S.I., W.M., Y.Z., M.A., Z.A., M.S., R.Y., A.J., G.T., J.C., H.L., Y.J., S.Z., X.H., J.R., Y.Q., Q.L., Z.Y., S.B., M.D., S.A., N.R., R.M., A.U.S., S.R., N.S., S.A., C.F., N.H., B.B., and M.L. contributed to the DH population seed bulk up and field trial work (planting, monitoring, onsite phenotyping, harvesting) and record the flowering data.