Author’s contributions: B.L., R.T.J, X.C and S.Q designed the study; S.Q and J.H performed the experiment; S.Q. and S.W. performed the analyses; S.Q, S.L, M.N., R.T.J and B.L. wrote the first manuscript draft, and all authors reviewed the manuscript and gave final approval for publication.Data availability statement: mitted to Dryad at https://datadrya